Astragalus 10a near Kazbegi, Georgia 170419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 10b near Kazbegi, Georgia 170419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 10c near Kazbegi, Georgia 170419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 10d near Kazbegi, Georgia 170419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 12a Cerro de la Cruz, Zapallar, Chile 171105 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 12b Cerro de la Cruz, Zapallar, Chile 171105 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 12c Cerro de la Cruz, Zapallar, Chile 171105 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 12e Cerro de la Cruz, Zapallar, Chile 171105 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 13b Pantallica area, Sicily 190417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 13c Pantallica area, Sicily 190417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 13d Pantallica area, Sicily 190417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 13e Pantallica area, Sicily 190417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 13f Pantallica area, Sicily 190417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 13g Pantallica area, Sicily 190417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 14a Napa Hai, Yunnan 110709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 14b Napa Hai, Yunnan 110709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 1h Dolna Kula, Bulgaria 120509 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 1i Dolna Kula, Bulgaria 120509 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 2a NW of Seki, Turkey 090410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 2b NW of Seki, Turkey 090410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 4c Shika, Yunnan 110712 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 4d Shika, Yunnan 110712 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 4e Shika, Yunnan 110712 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 4g Shika, Yunnan 110712 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 9a Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 9b Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 9c Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus acaulis 1 Baimashan, Yunnan 110710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus acaulis 2 Baimashan, Yunnan 110710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus acaulis 3 Baimashan, Yunnan 110710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus acaulis 4 Baimashan, Yunnan 110710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus acaulis 5 Baimashan, Yunnan 110710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus acaulis 6 Baimashan, Yunnan 110710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus acaulis 7 Baimashan, Yunnan 110710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus alopecuroides 3 Copenhagen Bot Garden 080710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus alopecuroides 4 Copenhagen Bot Garden 080710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus alopecuroides 5 Copenhagen Bot Garden 080710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus alpinus ssp alpinus 1 Söndre Knutshö, Dovre, Norway 820802 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus alpinus ssp alpinus 4 Långstrandberget near Ammerån, Jämtland 090702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus alpinus ssp alpinus 5 Långstrandberget near Ammerån, Jämtland 090702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus alpinus ssp alpinus 6 Långstrandberget near Ammerån, Jämtland 090702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus alpinus ssp arcticus 1 Unna Titer, Lule Lappmark 870717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus alpinus ssp arcticus 3 Unna Titer, Lule Lappmark 870717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus alpinus ssp arcticus 5 Kermesaari FI 120731 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus alpinus ssp arcticus 6 Kermesaari FI 120731 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus alpinus ssp arcticus 9 Kermesaari FI 120731 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus arenarius 1 Merkine, Lithuania 100714 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus arenarius 10 Vitemölla, Vitaby Sk 220609 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus arenarius 11 Vitemölla, Vitaby Sk 220609 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus arenarius 13 Vitemölla, Vitaby Sk 220609 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus arenarius 14 Vitemölla, Vitaby Sk 220609 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus arenarius 2 Merkine, Lithuania 100714 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus arenarius 3 between Kivik and Vitemölla, Vitaby Sk 080603 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus arenarius 4 between Kivik and Vitemölla, Vitaby Sk 080603 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus arenarius 5 between Kivik and Vitemölla, Vitaby Sk 080603 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus arenarius 6 between Kivik and Vitemölla, Vitaby Sk 080603 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus arenarius 7 between Kivik and Vitemölla, Vitaby, Skåne 790622 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus arenarius 8 between Kivik and Vitemölla, Vitaby, Skåne 790622 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus arenarius 9 between Kivik and Vitemölla, Vitaby, Skåne 920607 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus austro-aegaeus Othos, Karpathos, Greece 040521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus balearicus 1 Bocquer Valley, Mallorca 110420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus balearicus 2 Bocquer Valley, Mallorca 110420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus balearicus 3 Bocquer Valley, Mallorca 110420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus balearicus 4 Bocquer Valley, Mallorca 110420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus balearicus 5 Bocquer Valley, Mallorca 110420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus balearicus 6 Bocquer Valley, Mallorca 110420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus balearicus 7 Bocquer Valley, Mallorca 110420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus balearicus 8 Bocquer Valley, Mallorca 110420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus balearicus 9 Bocquer Valley, Mallorca 110420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus balfourianus 1 near Lijiang, Yunnan 110707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus balfourianus 3 Shangri-la, Yunnan 110712 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus boeticus 1 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus boeticus 2 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf armatus 1 Djebel Touila, Tunisia 820103 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf australis 1 Col d'Izoard FR 150709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf australis 2 Col d'Izoard FR 150709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf austriacus 1 Turda, Romania 090614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf austriacus 2 Turda, Romania 090614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf austriacus 6 Turda, Romania 090614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf cancellatus 2 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf lusitanicus 1 near Trujillo SP 140510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf lusitanicus 2 near Trujillo SP 140510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf monspessulanus 10 Sicily 190419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf monspessulanus 13 Sicily 190419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf monspessulanus 14 Sicily 190418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf monspessulanus 6 Sicily 190418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf monspessulanus 7 Sicily 190418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf monspessulanus 8 Sicily 190418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf spinosus 1 (cult) Brunnby Sk 130615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf spinosus 2 (cult) Brunnby Sk 130615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf spinosus 3 (cult) Brunnby Sk 130615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf turolensis 3 Hoces del Rio Duratón near Sepúlveda SP 140504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cicer 10 Lund Bot Garden 130816 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cicer 17 Les Freaux near La Meije FR 150706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cicer 8 Lund Bot Garden 130816 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus contortuplicatus 3 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus danicus 1 Klintehamn, Klinte, Gotland 070606 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus danicus 2 Klintehamn, Klinte, Gotland 070606 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus danicus 3 Klintehamn, Klinte, Gotland 070606 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus danicus 4 Klintehamn, Klinte, Gotland 070606 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus danicus 5 Klintehamn, Klinte, Gotland 070606 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus danicus 7 Klintehamn, Klinte, Gotland 070606 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus danicus 8 Peberholm DK 090804 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 1 Gorman Hills, CA 130409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 10 Antelope Valley Poppy Reserve, CA 130409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 15 Cabrillo National Monument, San Diego, California 130418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 16 San Diego, California 130417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 17 San Diego, California 130417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 2 Gorman Hills, CA 130409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 20 San Diego, California 130417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 21 San Diego, California 130417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 22 San Diego, California 130417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 23 San Diego, California 130417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 3 Gorman Hills, CA 130409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 4 Gorman Hills, CA 130409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 5 Gorman Hills, CA 130409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 9 Antelope Valley Poppy Reserve, CA 130409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus exscapus 1 (cult) Brunnby Sk 110521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus exscapus 10 Belsőbáránd, Hungary 240505 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus exscapus 11 Belsőbáránd, Hungary 240505 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus exscapus 2 (cult) Brunnby Sk 110521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus exscapus 3 (cult) Brunnby Sk 110521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus exscapus 4 (cult) Brunnby Sk 110521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus exscapus 5 (cult) Brunnby Sk 110521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus exscapus 6 (cult) Brunnby Sk 110521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus exscapus 7 (cult) Brunnby Sk 110521 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus exscapus 8 Belsőbáránd, Hungary 240505 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus exscapus 9 Belsőbáránd, Hungary 240505 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus falcatus 7 Lund Bot Garden 150612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus falcatus 8 Lund Bot Garden 150612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus frigidus 2 Abisko, Torne Lappmark 110725 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus frigidus 3 Abisko, Torne Lappmark 110725 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus frigidus 7 Stubnerkogel, Austria 080702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glaux 11 Sierra de las Nieves, Andalucía Spain 220409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glaux 13 Sierra de las Nieves, Andalucía Spain 220409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glaux 14 Sierra de las Nieves, Andalucía Spain 220409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glaux 15 Sierra de las Nieves, Andalucía Spain 220409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glaux 16 Sierra de las Nieves, Andalucía Spain 220409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glaux 17 Torcal de Antequera, Andalucía Spain 220418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glaux 18 Torcal de Antequera, Andalucía Spain 220418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glaux 19 Torcal de Antequera, Andalucía Spain 220418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glaux 5 near Benaoján, Andalusia Sp 120325 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glaux 7 near Benaoján, Andalusia Sp 120325 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glaux 8 near Benaoján, Andalusia Sp 120325 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glycyphylloides 1 near Milea, old road to Grevena, N Greece 180628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glycyphylloides 4 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glycyphyllus 1 Höllviken Sk 100626 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glycyphyllus 3 Höllviken Sk 100703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glycyphyllus 4 Höllviken Sk 100703 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hamosus 1 Agia Galini, Crete 110331 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hamosus 2 Agia Galini, Crete 110331 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hamosus 3 Agia Galini, Crete 110331 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hamosus 4 Agia Galini, Crete 110331 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hamosus 5 Agia Galini, Crete 110331 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hamosus 7 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hamosus 8 Copenhagen Bot Garden 130821 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hypoglottis 1 Hoces del Rio Duratón near Sepúlveda SP 140504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hypoglottis 10 Hoces del Rio Duratón near Sepúlveda SP 140504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hypoglottis 11 Hoces del Rio Duratón near Sepúlveda SP 140504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hypoglottis 2 Hoces del Rio Duratón near Sepúlveda SP 140504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hypoglottis 3 Hoces del Rio Duratón near Sepúlveda SP 140504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hypoglottis 8 Hoces del Rio Duratón near Sepúlveda SP 140504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lentiginosus 12 Joshua Tree National Park, California 130416 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lentiginosus 13 Joshua Tree National Park, California 130416 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lentiginosus 16 Joshua Tree National Park, California 130416 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lentiginosus 17 Joshua Tree National Park, California 130416 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lentiginosus 18 Joshua Tree National Park, California 130416 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lentiginosus 19 Joshua Tree National Park, California 130416 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lentiginosus 20 Joshua Tree National Park, California 130416 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lentiginosus 3 Wind Wolves Preserve, CA 130412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lentiginosus 4 Wind Wolves Preserve, CA 130412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lentiginosus 5 Wind Wolves Preserve, CA 130412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lentiginosus 6 Wind Wolves Preserve, CA 130412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lentiginosus 7 Wind Wolves Preserve, CA 130412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lentiginosus 8 Wind Wolves Preserve, CA 130412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lentiginosus 9 Wind Wolves Preserve, CA 130412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus longidentatus 1 Rambla de Tabernas, Andalucía Spain 220415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus longidentatus 2 Rambla de Tabernas, Andalucía Spain 220415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus longidentatus 3 Rambla de Tabernas, Andalucía Spain 220415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus longidentatus 4 Rambla de Tabernas, Andalucía Spain 220415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lusitanicus 10 S of Ojén, Andalusia SP 120326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lusitanicus 11 S of Ojén, Andalusia SP 120326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lusitanicus 12 S of Ojén, Andalusia SP 120326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lusitanicus 9 S of Ojén, Andalusia SP 120326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus massiliensis 2 Copenhagen Bot Garden 080513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus massiliensis 3 Copenhagen Bot Garden 080513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus mayeri 2 Devas near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus mayeri 3 Devas near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus mayeri 4 Devas near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus monspessulanus 1 Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus monspessulanus 10 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus monspessulanus 11 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus monspessulanus 12 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus monspessulanus 2 Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus monspessulanus 3 Copenhagen Bot Garden 140515 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus monspessulanus 4 Copenhagen Bot Garden 080513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus monspessulanus 5 Copenhagen Bot Garden 080513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus monspessulanus 6 Copenhagen Bot Garden 080513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus monspessulanus 7 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus monspessulanus 8 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus monspessulanus 9 Bergianska 150527 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus norvegicus 1 Unna Titer, Lule Lappmark 870714 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus onobrychis 8 Vashlovani Nat. Park, Georgia 170424 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus onobrychis 9 Vashlovani Nat. Park, Georgia 170424 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus oxyphysus 10 Carrizo Plain National Monument, CA 130412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus oxyphysus 11 Carrizo Plain National Monument, CA 130412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus oxyphysus 12 Carrizo Plain National Monument, CA 130412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus oxyphysus 6 Carrizo Plain National Monument, CA 130412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus oxyphysus 7 Carrizo Plain National Monument, CA 130412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus oxyphysus 8 Carrizo Plain National Monument, CA 130412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus oxyphysus 9 Carrizo Plain National Monument, CA 130412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus pachypus 1 Anza Borrego, California 130417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus pachypus 2 Anza Borrego, California 130417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus pachypus 3 Anza Borrego, California 130417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus pachypus 4 Anza Borrego, California 130417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus pachypus 5 Anza Borrego, California 130417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus pachypus 7 Anza Borrego, California 130417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus parnassi 1 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus parnassi 2 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus parnassi 3 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus parnassi 4 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus parnassi 5 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus pastellianus 1 Suatu, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus pelecinus 1 Barranco Angustias, La Palma 090502 - Photo: Richard Åkesson (© 2014 Jan Thomas Johansson).

Astragalus pelecinus 2 Petra, Lesvos, Greece 100516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus penduliflorus 2 E of Stugun church, Jämtland 090702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus penduliflorus 8 E of Stugun church, Jämtland 090702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus penduliflorus 9 E of Stugun church, Jämtland 090702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus persoonii 3 Algodones Dunes, CA 130420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus persoonii 4 Algodones Dunes, CA 130420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus persoonii 5 Algodones Dunes, CA 130420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus persoonii 6 Algodones Dunes, CA 130420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus pinetorum 1 Vashlovani Nat. Park, Georgia 170424 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus pinetorum 2 Vashlovani Nat. Park, Georgia 170424 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus pinetorum 3 Vashlovani Nat. Park, Georgia 170424 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus pinetorum 4 Vashlovani Nat. Park, Georgia 170424 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus trichopodus var lonchus 6 Torrey Pines, San Diego, CA 130418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus trichopodus var lonchus 7 Torrey Pines, San Diego, CA 130418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus yunnanensis 1 Baimashan, Yunnan 110710 - Photo: Magnus Thorell (© 2014 Jan Thomas Johansson).

Astragalus yunnanensis 2 Baimashan, Yunnan 110710 - Photo: Magnus Thorell (© 2014 Jan Thomas Johansson).

Astragalus yunnanensis 3 Baimashan, Yunnan 110710 - Photo: Magnus Thorell (© 2014 Jan Thomas Johansson).

Astragalus yunnanensis 4 Baimashan, Yunnan 110710 - Photo: Magnus Thorell (© 2014 Jan Thomas Johansson).

Astragalus 12d Cerro de la Cruz, Zapallar, Chile 171105 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 13a Pantallica area, Sicily 190417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 1a Dolna Kula, Bulgaria 120509 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 1b Dolna Kula, Bulgaria 120509 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 1c Dolna Kula, Bulgaria 120509 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 1d Dolna Kula, Bulgaria 120509 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 1e Dolna Kula, Bulgaria 120509 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 1f Dolna Kula, Bulgaria 120509 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 1g Dolna Kula, Bulgaria 120509 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 2c NW of Seki, Turkey 090410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 2d NW of Seki, Turkey 090410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 2e NW of Seki, Turkey 090410 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 3a Baimashan, Yunnan 110710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 3b Baimashan, Yunnan 110710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 4a Shika, Yunnan 110712 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 4b Shika, Yunnan 110712 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 4f Shika, Yunnan 110712 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 9d Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus 9e Copenhagen Bot Garden 130516 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus alopecuroides 1 Copenhagen Bot Garden 080710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus alopecuroides 2 Copenhagen Bot Garden 080710 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus alpinus ssp alpinus 2 Söndre Knutshö, Dovre, Norway 820802 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus alpinus ssp alpinus 3 S of Stugun, Jämtland 870718 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus alpinus ssp arcticus 2 Unna Titer, Lule Lappmark 870717 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus alpinus ssp arcticus 4 Kermesaari FI 120731 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus alpinus ssp arcticus 7 Kermesaari FI 120731 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus alpinus ssp arcticus 8 Kermesaari FI 120731 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus arenarius 12 Vitemölla, Vitaby Sk 220609 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus asper 1 Belsőbáránd, Hungary 240505 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus asper 2 Belsőbáránd, Hungary 240505 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus austriacus 1 Belsőbáránd, Hungary 240505 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus austriacus 2 Belsőbáránd, Hungary 240505 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus austriacus 3 Belsőbáránd, Hungary 240505 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus austriacus 4 Belsőbáránd, Hungary 240505 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus austriacus 5 Belsőbáránd, Hungary 240505 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus balfourianus 2 near Lijiang, Yunnan 110707 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus balfourianus 4 Shangri-la, Yunnan 110712 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus balfourianus 5 Shangri-la, Yunnan 110712 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf armatus 2 Djebel Touila, Tunisia 820103 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf austriacus 3 Turda, Romania 090614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf austriacus 4 Turda, Romania 090614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf austriacus 5 Turda, Romania 090614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf austriacus 7 Turda, Romania 090614 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf cancellatus 1 Yeghegnadzan mountains, Armenia 180511 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf lusitanicus 3 near Trujillo SP 140510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf lusitanicus 4 near Trujillo SP 140510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf lusitanicus 5 near Trujillo SP 140510 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf monspessulanus 1 Monte Aragón, N Spain 180526 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf monspessulanus 11 Sicily 190419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf monspessulanus 12 Sicily 190419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf monspessulanus 2 Monte Aragón, N Spain 180526 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf monspessulanus 3 Monte Aragón, N Spain 180526 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf monspessulanus 4 Monte Aragón, N Spain 180526 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf monspessulanus 5 Sicily 190419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf monspessulanus 9 Sicily 190419 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf onobrychis 1 NW of Dadia, Thracia, Greece 060606 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf onobrychis 3 NW of Dadia, Thracia, Greece 060606 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf onobrychis 4 NW of Dadia, Thracia, Greece 060606 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf turolensis 1 Hoces del Rio Duratón near Sepúlveda SP 140504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cf turolensis 2 Hoces del Rio Duratón near Sepúlveda SP 140504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cicer 1 E of Veseli, Czekia 950630 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cicer 11 Lund Bot Garden 130816 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cicer 12 Bergianska, Stockholm 010824 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cicer 13 Les Freaux near La Meije FR 150706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cicer 14 Les Freaux near La Meije FR 150706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cicer 15 Les Freaux near La Meije FR 150706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cicer 16 Les Freaux near La Meije FR 150706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cicer 2 Lund Bot Garden 100706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cicer 3 Lund Bot Garden 100706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cicer 4 Lund Bot Garden 100706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cicer 5 Lund Bot Garden 100706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cicer 6 Lund Bot Garden 100706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cicer 7 Lund Bot Garden 100706 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus cicer 9 Lund Bot Garden 130816 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus contortuplicatus 1 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus contortuplicatus 2 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus contortuplicatus 4 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus contortuplicatus 5 Copenhagen Bot Garden 140719 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus danicus 6 Klintehamn, Klinte, Gotland 070606 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 11 Antelope Valley Poppy Reserve, CA 130409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 12 Cabrillo National Monument, San Diego, California 130418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 13 Cabrillo National Monument, San Diego, California 130418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 14 Cabrillo National Monument, San Diego, California 130418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 18 San Diego, California 130417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 19 San Diego, California 130417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 24 San Diego, California 130417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 6 Antelope Valley Poppy Reserve, CA 130409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 7 Antelope Valley Poppy Reserve, CA 130409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus douglasii 8 Antelope Valley Poppy Reserve, CA 130409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus falcatus 1 (cult) Brunnby Sk (orig Bergianska, Stockholm) 130615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus falcatus 10 Lund Bot Garden 150612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus falcatus 11 Lund Bot Garden 150612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus falcatus 12 Lund Bot Garden 150612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus falcatus 2 (cult) Brunnby Sk (orig Bergianska, Stockholm) 130615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus falcatus 3 (cult) Brunnby Sk (orig Bergianska, Stockholm) 130615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus falcatus 4 (cult) Brunnby Sk (orig Bergianska, Stockholm) 130615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus falcatus 5 (cult) Brunnby Sk (orig Bergianska, Stockholm) 130615 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus falcatus 6 Bergianska, Stockholm 010824 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus falcatus 9 Lund Bot Garden 150612 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus frigidus 1 Abisko, Torne Lappmark 110725 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus frigidus 4 Söndre Knutshö, Dovre, Norway 820802 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus frigidus 5 Söndre Knutshö, Dovre, Norway 820802 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus frigidus 6 Stubnerkogel, Austria 080702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glaux 1 near Benaoján, Andalusia Sp 120325 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glaux 10 Sierra de las Nieves, Andalucía Spain 220409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glaux 12 Sierra de las Nieves, Andalucía Spain 220409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glaux 2 near Benaoján, Andalusia Sp 120325 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glaux 3 near Benaoján, Andalusia Sp 120325 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glaux 4 near Benaoján, Andalusia Sp 120325 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glaux 6 near Benaoján, Andalusia Sp 120325 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glaux 9 Sierra de las Nieves, Andalucía Spain 220409 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glycyphylloides 1 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glycyphylloides 2 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glycyphylloides 2 near Milea, old road to Grevena, N Greece 180628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glycyphylloides 3 Copenhagen Bot Garden 130625 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glycyphylloides 3 near Milea, old road to Grevena, N Greece 180628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glycyphylloides 4 near Milea, old road to Grevena, N Greece 180628 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glycyphyllus 2 Höllviken Sk 100626 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glycyphyllus 5 Stammershalle, Bornholm DK 120602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glycyphyllus 6 Stammershalle, Bornholm DK 120602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glycyphyllus 7 Stammershalle, Bornholm DK 120602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus glycyphyllus 8 Stammershalle, Bornholm DK 120602 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus gummifer Bergianska, Stockholm 050913 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hamosus 6 Agia Galini, Crete 110331 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hypoglottis 5 Hoces del Rio Duratón near Sepúlveda SP 140504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hypoglottis 6 Hoces del Rio Duratón near Sepúlveda SP 140504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hypoglottis 7 Hoces del Rio Duratón near Sepúlveda SP 140504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hypoglottis 9 Hoces del Rio Duratón near Sepúlveda SP 140504 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hypoglottis ssp hypoglottis 1 N of La Vachette FR 150709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hypoglottis ssp hypoglottis 10 Le Fontenil N of Briancon FR 150709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hypoglottis ssp hypoglottis 11 Le Fontenil N of Briancon FR 150709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hypoglottis ssp hypoglottis 2 N of La Vachette FR 150709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hypoglottis ssp hypoglottis 3 N of La Vachette FR 150709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hypoglottis ssp hypoglottis 4 N of La Vachette FR 150709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hypoglottis ssp hypoglottis 5 N of La Vachette FR 150709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hypoglottis ssp hypoglottis 6 Le Fontenil N of Briancon FR 150709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hypoglottis ssp hypoglottis 7 Le Fontenil N of Briancon FR 150709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hypoglottis ssp hypoglottis 8 Le Fontenil N of Briancon FR 150709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus hypoglottis ssp hypoglottis 9 Le Fontenil N of Briancon FR 150709 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus laxmannii 1 Lund Bot Garden 140617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus laxmannii 2 Lund Bot Garden 140617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus laxmannii 3 Lund Bot Garden 140617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus laxmannii 4 Lund Bot Garden 140617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus laxmannii 5 Lund Bot Garden 140617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus laxmannii 6 Lund Bot Garden 140617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus laxmannii 7 Lund Bot Garden 140617 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lentiginosus 1 Wind Wolves Preserve, CA 130412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lentiginosus 10 Wind Wolves Preserve, CA 130412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lentiginosus 11 Wind Wolves Preserve, CA 130412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lentiginosus 14 Joshua Tree National Park, California 130416 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lentiginosus 15 Joshua Tree National Park, California 130416 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lentiginosus 2 Wind Wolves Preserve, CA 130412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus longidentatus 5 Rambla de Tabernas, Andalucía Spain 220415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus longidentatus 6 Rambla de Tabernas, Andalucía Spain 220415 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lusitanicus 1 S of Ojén, Andalusia SP 120326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lusitanicus 13 S of Ojén, Andalusia SP 120326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lusitanicus 14 S of Ojén, Andalusia SP 120326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lusitanicus 2 S of Ojén, Andalusia SP 120326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lusitanicus 3 S of Ojén, Andalusia SP 120326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lusitanicus 4 S of Ojén, Andalusia SP 120326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lusitanicus 5 S of Ojén, Andalusia SP 120326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lusitanicus 6 S of Ojén, Andalusia SP 120326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lusitanicus 7 S of Ojén, Andalusia SP 120326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus lusitanicus 8 S of Ojén, Andalusia SP 120326 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus massiliensis 1 Copenhagen Bot Garden 080513 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus mayeri 1 Devas near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus mayeri 5 Devas near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus mayeri 6 Devas near Lake Mikri Prespa, N Greece 180629 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus mongolicus 1 Bergianska, Stockholm 050621 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus mongolicus 2 Bergianska, Stockholm 050621 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus norvegicus 2 Unna Titer, Lule Lappmark 870714 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus norvegicus 3 Unna Titer, Lule Lappmark 870714 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus onobrychis 1 Bergianska, Stockholm 030606 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus onobrychis 2 Bergianska, Stockholm 030606 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus onobrychis 3 Vashlovani Nat. Park, Georgia 170424 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus onobrychis 4 Vashlovani Nat. Park, Georgia 170424 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus onobrychis 5 Vashlovani Nat. Park, Georgia 170424 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus onobrychis 6 Vashlovani Nat. Park, Georgia 170424 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus onobrychis 7 Vashlovani Nat. Park, Georgia 170424 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus oxyphysus 1 Carrizo Plain National Monument, CA 130412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus oxyphysus 2 Carrizo Plain National Monument, CA 130412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus oxyphysus 3 Carrizo Plain National Monument, CA 130412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus oxyphysus 4 Carrizo Plain National Monument, CA 130412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus oxyphysus 5 Carrizo Plain National Monument, CA 130412 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus pachypus 6 Anza Borrego, California 130417 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus pastellianus 2 Suatu, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus pastellianus 3 Suatu, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus pastellianus 4 Suatu, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus pastellianus 5 Suatu, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus pastellianus 6 Suatu, Romania 090613 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus penduliflorus 1 E of Stugun church, Jämtland 090702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus penduliflorus 10 E of Stugun church, Jämtland 090702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus penduliflorus 11 E of Stugun church, Jämtland 090702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus penduliflorus 12 N of Gagnef stn, Dalarne 930723 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus penduliflorus 13 N of Gagnef stn, Dalarne 930723 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus penduliflorus 3 E of Stugun church, Jämtland 090702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus penduliflorus 4 E of Stugun church, Jämtland 090702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus penduliflorus 5 E of Stugun church, Jämtland 090702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus penduliflorus 6 E of Stugun church, Jämtland 090702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus penduliflorus 7 E of Stugun church, Jämtland 090702 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus persoonii 1 Algodones Dunes, CA 130420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus persoonii 2 Algodones Dunes, CA 130420 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus trichopodus var lonchus 1 Torrey Pines, San Diego, CA 130418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus trichopodus var lonchus 2 Torrey Pines, San Diego, CA 130418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus trichopodus var lonchus 3 Torrey Pines, San Diego, CA 130418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus trichopodus var lonchus 4 Torrey Pines, San Diego, CA 130418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).

Astragalus trichopodus var lonchus 5 Torrey Pines, San Diego, CA 130418 - Photo: Jan Thomas Johansson (© 2014 Jan Thomas Johansson).